In Paris, Jacques Saunière is pursued through the Louvre's Grand Gallery by albino monk Silas (Paul Bettany), demanding the Priory's clef de voûte or "keystone." Saunière confesses the keystone is kept in the sacristy of Church of Saint-Sulpice "beneath the Rose" before Silas shoots him. At the American University of Paris, Robert Langdon, a symbologist who is a guest lecturer on symbols and the sacred feminine, is summoned to the Louvre to view the crime scene. He discovers the dying Saunière has created an intricate display using black light ink and his own body and blood. Captain Bezu Fache (Reno) asks him for his interpretation of the puzzling scene.
Silas calls a mysterious man known as "The Teacher", revealing that he has killed all four protectors of the keystone and that all confirmed the same location. He dons a metal cilice on his thigh and proceeds to flagellate himself with a whip for the sins of murder. Facilitated by Bishop Manuel Aringarosa, Silas then travels to Saint-Sulpice and is admitted by an elderly nun; left alone, he excavates beneath the floor of the church to find a stone saying only JOB 38:11. He confronts the nun, who quotes the passage: "Hitherto shalt thou come, but no further." Realizing that he has been deceived, Silas is enraged and kills the nun.
Sophie Neveu, a cryptologist with the French police,and the granddaughter of Saunière, enters the Louvre as well and slips Langdon a message which leads him to the bathroom. There, Sophie meets him and tells him that he is being tracked, a GPS tracking dot has been slipped into his jacket and that he is a primary suspect in the murder case because of a line of text found by the corpse. Sophie, however, believes that Saunière wanted to pass a hidden message on to her and that he had wanted to bring Langdon into the equation so that he could help her crack the code.
Buying some time by throwing the tracking device into the back of a truck, the pair begins exploring the Louvre, finding more anagram messages that Saunière had left behind. Many of these relate to Leonardo da Vinci's art, and the pair find a key with a Fleur-de-lis behind Madonna of the Rocks. Langdon deduces from this that Saunière was a member of the Priory of Sion, a secret society associated with the Knights Templar. At this point, Langdon also asserts a connection with the Holy Grail.
Pursued by the French police and cut off from the United States Embassy, the pair escape to the Bois de Boulogne where Langdon closely inspects the key. He notices an inscription on the side – an address. The address directs them to the Depository Bank of Zurich where the key is used for a safety deposit box.
In the bank, they find Saunière's deposit box and open it using the 10 digit Fibonacci numbers in order (1123581321). Inside the box, they find a rosewood container, which contains a cryptex: a cylindrical container with five alphabetical dials which must be arranged in the correct sequence to spell out a 5-letter code word, in order to open and access the papyrus message inside. Using force to open the cryptex would break a vial of vinegar inside, which would dissolve the papyrus and destroy the message.
Unfortunately, the police are called by a security guard and they are forced to leave. The bank manager, Andre Vernet, assists them in escaping by taking them as passengers in an armoured van to escape the routine checks of the police. Eventually, they come to a sudden stop and Vernet forces them at gunpoint to give him the cryptex. Langdon tricks Vernet and disarms him and he and Sophie escape with the cryptex in their hands.
Langdon suggests that they visit his friend, Leigh Teabing (Ian McKellen), for assistance to opening the cryptex. Leigh turns out to be an enthusiastic seeker of the Holy Grail, which he believes is not actually a cup but instead Mary Magdalene who, he claims, was pregnant at the time of Christ's crucifixion. Teabing tells Sophie that the Priory of Sion was formed to protect the descendants of Jesus. Jacques Saunière was believed to be a part of this society and Teabing suspects that he was training Sophie to join it. Silas, meanwhile, breaks into Teabing's mansion and attempts to steal the cryptex. Teabing uses his cane to knock Silas out and they escape again, taking the butler, Remy Jean, and Silas with them. The group escapes in Teabing's plane, following the next clue to London. Fache learns of their destination, and alerts the London Metropolitan Police to apprehend them at the airport. But Teabing manages to slip the party past the police with a trick of misdirection.
Teabing leads Langdon and Neveu to the Temple Church in London, which is shown to be a red herring. Silas is freed by Remy Jean, who is revealed to be a follower of The Teacher as well. The two take Teabing hostage, and Silas, believing Remy to be The Teacher, holes up in an Opus Dei safehouse. Remy is killed by the mysterious man after deceiving Silas. Silas accidentally shoots Aringarosa and is shot by the police. Aringarosa is taken to the hospital and apprehended by Fache.
Langdon is betrayed by Teabing, who is revealed to be the Teacher. He escapes with the cryptex, and attempts to find the next clue at Isaac Newton's tomb in Westminster Abbey, as Newton was a member of the Priory. Langdon and Neveu catch up with him, and chase him into the nearby chapter house. Teabing explains that he wanted to find Mary Magdalene's remains for two reasons; first, to prove that all of his research was right. Furthermore, Teabing claims that the Priory was charged with revealing the entire conspiracy back in 2000 and when they continued to conceal it, he became The Teacher to force the secret to the public eye. When Sophie admits she does not know the combination to the cryptex, Teabing threatens to shoot Sophie if Langdon does not open the cryptex for him. Langdon takes the cryptex for a moment to think of the answer, then throws the cryptex into the air, leaving Teabing to fumble and destroy it.
Distraught at not receiving the code, Teabing is arrested, but Langdon had cracked the code ('Apple', a reference to Newton) and removed the clue from the cryptex before destroying it. Using the clue, they travel to Rosslyn Chapel in Scotland where Magdalene's remains had previously been hidden. The remains had since been moved, but they meet other members of the secret organization that protected her, as well as finding scores of documentation chronicling the "Sang Real" ("Royal Blood") bloodline. Sophie is actually Magdalene's descendant and, therefore, of Jesus Christ. They vow to keep her safe before going their separate ways. Neveu and Langdon debate the significance of her heritage, and Langdon finally counsels her that it's up to her what to make of the legacy she has been given.
In Paris, Langdon accidentally cuts himself while shaving and the line of blood on the sink reminds him of the Rose Line. He follows the Rose Line and finds the location of the Holy Grail, buried under the pyramid in the Louvre. Langdon then kneels above Mary Magdalene's tomb as the Knights Templar did before him.
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